Parrotfish Teeth: Stiff Biominerals Whose Microstructure Makes Them Tough and Abrasion-Resistant to Bite Stony Corals

Matthew A. Marcus, Shahrouz Amini, Cayla A. Stifler, Chang Yu Sun, Nobumichi Tamura, Hans A. Bechtel, Dilworth Y. Parkinson, Harold S. Barnard, Xiyue X.X. Zhang, J. Q.Isaiah Chua, Ali Miserez, Pupa U.P.A. Gilbert

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42 Citations (Scopus)

Abstract

Parrotfish (Scaridae) feed by biting stony corals. To investigate how their teeth endure the associated contact stresses, we examine the chemical composition, nano- and microscale structure, and the mechanical properties of the steephead parrotfish Chlorurus microrhinos tooth. Its enameloid is a fluorapatite (Ca5(PO4)3F) biomineral with outstanding mechanical characteristics: the mean elastic modulus is 124 GPa, and the mean hardness near the biting surface is 7.3 GPa, making this one of the stiffest and hardest biominerals measured; the mean indentation yield strength is above 6 GPa, and the mean fracture toughness is ∼.5 MPa·m1/2, relatively high for a highly mineralized material. This combination of properties results in high abrasion resistance. Fluorapatite X-ray absorption spectroscopy exhibits linear dichroism at the Ca L-edge, an effect that makes peak intensities vary with crystal orientation, under linearly polarized X-ray illumination. This observation enables polarization-dependent imaging contrast mapping of apatite, a method to quantitatively measure and display nanocrystal orientations in large, pristine arrays of nano- and microcrystalline structures. Parrotfish enameloid consists of 100 nm-wide, microns long crystals co-oriented and assembled into bundles interwoven as the warp and the weave in fabric and therefore termed fibers here. These fibers gradually decrease in average diameter from 5 μm at the back to 2 μm at the tip of the tooth. Intriguingly, this size decrease is spatially correlated with an increase in hardness.

Original languageEnglish
Pages (from-to)11856-11865
Number of pages10
JournalACS nano
Volume11
Issue number12
DOIs
Publication statusPublished - 2017 Dec 26

All Science Journal Classification (ASJC) codes

  • General Materials Science
  • General Engineering
  • General Physics and Astronomy

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